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81 Spatial dynamics of host parasitoid systems In 6 we noted that Nicholson and Bailey had anticipated that spatial structure, in particular migration and local extinction, would lead to persistence in their model of host parasitoid interactions Comins et al (1992) used the Nicholson Bailey model to explore the possible outcomes of spatial dynamics In this study it was assumed that the host and parasitoid were distributed among a square grid of square cells or patches of width n These types of model systems, known collectively as cellular automata, have been widely used in both plant and animal studies to address a variety of issues including ecological stability and effects of invasive species (eg Crawley & May 1987, Silvertown et al 1992, Colasanti & Grime 1993, Huang et al 2008; see Wolfram 1984 for a mathematical overview) Comins et al.

The requirement is inapplicable if the inclusion of women and members of minority groups is inappropriate with respect to the health of subjects to the purpose of the research or under other designated circumstances The cost of inclusion is not a permissible factor in determining whether inclusion is inappropriate A sponsor or conductor of research, who would have selected all the subjects on some other basis, now must pay attention to race and gender and include enough women and minorities to make statistically meaningful assessments of drug safety and ef cacy for those groups as well Although no speci c race is singled out or numbers speci ed, the law does impose a requirement to include subjects on the basis of race and gender when subjects would have been chosen on the basis of inclusion criteria insensitive to the needs of persons in those groups.

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(1992) had two phases of dynamics: reproduction/parasitism and dispersal The former was modelled using the Nicholson Bailey model The latter had the following rules 1 A fraction of the hosts and parasitoids leave the patch (grid cell) and the remainder stay to reproduce in their patch 2 The fraction dispersing is equally divided between the eight neighbouring patches There is only one movement per generation Longer-range dispersal is excluded 3 There are re ective boundary conditions in which dispersing individuals are prevented from crossing the boundary and remain in the edge patch Thus there is an explicit edge effect in this model, in contrast to some other cellular automata models In small arenas of less than 10 cells by 10 cells extinction of host and parasitoid occurred within a few hundred generations of the simulations, underlining the inherent instability of the Nicholson Bailey model.

Without a mandatory inclusion requirement, too few representatives of each group might be included to provide meaningful information about them The racial and gender classi cations at issue here, however, are justi ed to achieve the government s goal of spending research funds in a way that will yield useful information about all citizens, including women and minorities, not simply about whites or males, as has been the case in the past Meeting the health care needs of all its citizens, including women and minorities, is a compelling state interest Because biological and genetic variation in women and minorities may differ signi cantly from that in white males, requiring their inclusion in funded research is essential if valid information about their health is also to be produced.

However, when the arena size was increased to between 15 and 30 patches, three general types of spatial dynamics were found which Comins et al described as spirals, spatial chaos and crystal lattices (Fig 81a) The key feature of the three dynamic types is that they permit long-term persistence of the host and parasitoid within a relatively narrow range of population densities (Fig 81b), as 135.

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Alternatives such as private investment by drug companies are not as likely to achieve that goal, for the minority subgroups involved may be too small to justify investment for that purpose Requiring inclusion of women and minorities in research is not based on any stereotype of the inferiority or inappropriateness of women and racial minorities, nor does it deny males or whites anything that they are due based on their merit or need Although motivated in part by the past exclusion of women and minorities from much biomedical research, the NIH program.

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